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Introduction to the Caryophyllids

the Betalain Bunch

The caryophyllids are a large group of eudicot flowering plants -- in fact they include more than 11,000 species, or about 7% of all flowering plants. Most of these species belong to one of three caryophyllid families: Aizoaceae (ice plants), Cactaceae (cacti), or Caryophyllaceae (carnations).

Most caryophyllids share a number of unusual characteristics in their pigmentation and their morphology. They contain betalain compounds, a group of redddish and yellow pigments that are not found in any other plants. If you have ever heard the expression "beet red", then you are aware of the strikingly red color that red betalains can create, since beets are members of this group.

Time bar

The Plantae includes all land plants: mosses, ferns, conifers, flowering plants, and so on - an amazing range of diverse forms. With more than 250,000 species, they are second in size only to the arthropoda.

Plants have been around for a very long time. The plants first appeared in the Ordovician, but did not begin to resemble modern plants until the Late Silurian. By the close of the Devonian, about 360 million years ago, there were a wide variety of shapes and sizes of plants around, including tiny creeping plants and tall forest trees.

The most striking, and important, feature of plants is their green color, the result of a pigment called chlorophyll. Plants use chlorophyll to capture light energy, which fuels the manufacture of food—sugar, starch, and other carbohydrates. Without these food sources, most life on earth would be impossible. There would still be mushrooms and algae, but there would be no fruits, vegetables, grains, or any animals (which ultimately rely on plants for their food too!)

Another important contribution of plants is their shaping of the environment. Think of a place without plants. The only such places on earth are the arctic wastelands, really arid deserts, and the deep ocean. Everywhere else, from the tundra to the rainforest to the desert, is populated by plants. In fact, when we think of a particular landscape, it is the plants which first come to mind. Try to picture a forest without trees, or a prairie without grasses. It is the plants which produce and maintain the terrestrial environment as we know it.

The Paleozoic is bracketed by two of the most important events in the history of animal life. At its beginning, multicelled animals underwent a dramatic "explosion" in diversity, and almost all living animal phyla appeared within a few millions of years. At the other end of the Paleozoic, the largest mass extinction in history wiped out approximately 90% of all marine animal species. The causes of both these events are still not fully understood and the subject of much research and controversy. Roughly halfway in between, animals, fungi, and plants alike colonized the land, the insects took to the air, and the limestone shown in this picture was deposited near Burlington, Missouri.

The Paleozoic took up over half of the Phanerozoic, approximately 300 million years. During the Paleozoic there were six major continental land masses; each of these consisted of different parts of the modern continents. For instance, at the beginning of the Paleozoic, today's western coast of North America ran east-west along the equator, while Africa was at the South Pole. These Paleozoic continents experienced tremendous mountain building along their margins, and numerous incursions and retreats of shallow seas across their interiors. Large limestone outcrops, like the one shown above, are evidence of these periodic incursions of continental seas.

Many Paleozoic rocks are economically important. For example, much of the limestone quarried for building and industrial purposes, as well as the coal deposits of western Europe and the eastern United States, were formed during the Paleozoic. 

Phanerozoic Eon
(544 mya to present)
Cenozoic Era
(65 mya to today)
Quaternary (1.8 mya to today)
       Holocene (11,000 years to today)
       Pleistocene (1.8 mya to 11,000 yrs)
Tertiary (65 to 1.8 mya)
       Pliocene (5 to 1.8 mya)
       Miocene (23 to 5 mya)
       Oligocene (38 to 23 mya)
       Eocene (54 to 38 mya)
       Paleocene (65 to 54 mya)
 
Mesozoic Era
(245 to 65 mya)
Cretaceous (146 to 65 mya)
Jurassic (208 to 146 mya)
Triassic (245 to 208 mya)
 
Paleozoic Era
(544 to 245 mya)
Permian (286 to 245 mya)
Carboniferous (360 to 286 mya)
       Pennsylvanian (325 to 286 mya)
       Mississippian (360 to 325 mya)

Devonian (410 to 360 mya)
Silurian (440 to 410 mya)
Ordovician (505 to 440 mya)
Cambrian (544 to 505 mya)
        Tommotian (530 to 527 mya)
 
Precambrian Time
(4,500 to 544 mya)
Proterozoic Era
(2500 to 544 mya)
Neoproterozoic (900 to 544 mya)
       Vendian (650 to 544 mya)
Mesoproterozoic (1600 to 900 mya)
Paleoproterozoic (2500 to 1600 mya)
 
Archaean
(3800 to 2500 mya)
 
Hadean
(4500 to 3800 mya)

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